Publications

Adrain, J.M. (2026) The trilobites Smoothcanyonops gen. nov. and Ceeops gen. nov. from the Early Ordovician (mid-Tremadocian; Stairsian) of the Great Basin, western USA. Bulletins of Geoscience, 101, 49–64. Link (open access)

Adrain, J.M. & Pérez-Peris, F. (2026) The Early Ordovician (Floian) bathyurid trilobite Jimlochaspis n. gen., with species from western Utah and Greenland. Palaeoworld, 35, 201110.

Adrain, J.M., Westrop, S.R., Landing, E., Karim, T.S., Losso, S.R., Ng, R.Y., Bradley, A.B. & Pérez-Peris, F. (2025) Trilobite biostratigraphy of the upper Skullrockian Stage (Lower Ordovician; lower Tremadocian) on the northern Laurentian margin, western United States. Australasian Palaeontological Memoirs, 57, 397–432.

Adrain, J.M. (2025) The Early Ordovician bathyurid trilobites Licnocephala and Ibexocephala. Journal of Paleontology. Link (open access)

Pérez-Peris, F., Adrain, J.M. & Daley, A.C. (2024) Phylogenetics and systematics of the subfamilies Cheirurinae and Deiphoninae (Trilobita). Swiss Journal of Palaeontology, 143, 43. Link (open access)

Pérez-Peris, F., Adrain, J.M. & Allison C. Daley (2024) Ordovician paleobiogeography of the Suborder Cheirurina (Trilobita). Palaeogeography, Palaeoclimatology, Palaeoecology, 647, 112222. Link (open access)

Westrop, S.R., Welch, K.F., Engel, M.H. & Adrain, J.M. (2023) Faunal and paleoenvironmental changes at a Cambrian (Jiangshanian; Steptoean–Sunwaptan boundary interval) trilobite extinction event, in contrasting deep and shallow subtidal settings, Nevada and Oklahoma. Journal of Paleontology. Link (open access)

Adrain, J.M. & Pérez-Peris, F. (2023) Funeralaspis n. gen.: a new odontopleurine trilobite from the early Middle Ordovician (Dapingian) of Death Valley, eastern California, USA, and the classification of Ordovician odontopleurines. Zootaxa, 5336, 509–529.


2022 Oenonella
Adrain, J.M. & Fortey, R.A. (2022) The Ordovician trilobite Oenonella and the new family Oenonellidae, with new species from western Newfoundland, Canada. Journal of Paleontology. Link (open access)



Forteyaspis
Karim, T.S. & Adrain, J.M. (2022) The phylogenetic affinity of the Ordovician trilobites Agerina, Forteyaspis gen. nov., and related genera, with new and revised species from Canada and the United States. Canadian Journal of Earth Sciences, 59, 156–179.



2021TCCheirurids
Adrain, J.M. & Pérez-Peris, F. (2021) Middle Ordovician (Darriwilian) cheirurid trilobites from the Table Cove Formation, western Newfoundland, Canada. Zootaxa, 5041, 1–73.



2021 Gonioteloides
Adrain, J.M. & Karim, T.S. (2021) Systematics of the Early Ordovician (late Tremadocian; Stairsian) trilobite Gonioteloides Kobayashi, with species from the Great Basin, western USA. Journal of Paleontology, 95, 1022–1047. Link (open access)


HughesNormative
Hughes, N.C., Adrain, J.M., Holmes, J.D., Hong, P.S., Hopkins, M.J., Hou, J.-B., Minelli, A., Park, T.-Y. S., Paterson, J.R., Peng, J., Webster, M., Zhang, X.-G., Zhang, X.-L. & Fusco, G. (2020) Articulated trilobite ontogeny: suggestions for a methodological standard. Journal of Paleontology, 95, 298–304. Link (open access)


2020 Atlanticalymene small pub
Adrain, J.M., Karim, T.S. & McAdams, N.E.B. (2020) Atlanticalymene, a new genus of Middle Ordovician (Darriwilian) calymenine trilobites, and revision of the calymenoidean genus Protocalymene Ross. Zootaxa, 4859, 1–55.


2018 Tesselacauda small pub
Adrain, J.M. & Karim, T.S. (2019) Revision of the Early Ordovician (late Tremadocian; Stairsian) cheirurid trilobite Tesselacauda Ross, with species from the Great Basin, western USA. Zootaxa, 4661, 201–255.


2018 Ibexaspis small pub
McAdams, N.E.B., Adrain, J.M. & Karim, T.S. (2018) The pliomerid trilobite Ibexaspis and related new genera, with species from the Early Ordovician (Floian; Tulean, Blackhillsian) of the Great Basin, western USA. Zootaxa, 4525, 1–152.


2016 Adrain & Westrop taph small pub
Adrain, J.M. & Westrop, S.R. (2016) Testing for taphonomic bias in deep time using trilobite sclerite ratios. Palaeogeography, Palaeoclimatology, Palaeoecology, 463, 205–215.


2016 Irvingella small pub
Westrop, S.R. & Adrain, J.M. (2016) Revision of Irvingella tropica Öpik, 1963 from Australia and related species from North America: implications for correlation of the base of the Jiangshanian Stage (Cambrian, Furongian). Memoirs of the Association of Australasian Palaeontologists, 49, 395–432.


Saltzman et al 2015 copy
Saltzman, M.R., Edwards, C.T., Adrain, J.M. & Westrop, S.R. (2015) Persistent oceanic anoxia and elevated extinction rates separate the Cambrian and Ordovician radiations. Geology, 43, 807–810.


2014 Pseudohystricurus
Adrain, J.M., Karim, T.S. & Westrop, S.R. (2014) The Early Ordovician (upper Tremadocian; Stairsian) dimeropygid trilobite Pseudohystricurus Ross. Memoirs of the Association of Australasian Palaeontologists, 45, 215–232.


2014 Bathyurina

Adrain, J.M., Karim, T.S. & Westrop, S.R. (2014) The Early Ordovician (Floian) bathyurid trilobite genera Jeffersonia, Cullisonia, and Bathyurina. Memoirs of the Association of Australasian Palaeontologists, 45, 305–347.


2014 Parakoldinioidia
Westrop, S.R. & Adrain, J.M. (2014) The missisquoiid trilobite Parakoldinioidia Endo, 1937, in the latest Cambrian of Oklahoma and Texas, and its biostratigraphic significance. Memoirs of the Association of Australasian Palaeontologists, 45, 117–152.


2014 Stairsian
Adrain, J.M., Karim, T.S., Westrop, S.R., & Landing, E. (2014) Trilobite biostratigraphy of the Stairsian Stage (upper Tremadocian) of the Ibexian Series, Lower Ordovician, western United States. Memoirs of the Association of Australasian Palaeontologists, 45, 167–214.


2013 Geol Soc
Adrain, J.M. (2013) A synopsis of Ordovician trilobite distribution and diversity. In: Harper, D.A.T. & Servais, T. (Eds.), Early Palaeozoic Palaeobiogeography and Palaeogeography. Geological Society, London, Memoirs, 38, 293–332.



2013 Biceragnotus

Westrop, S.R. & Adrain, J.M. (2013) Biogeographic shifts in a transgressive succession: The Cambrian (Furongian; Jiangshanian; latest Steptoean–earliest Sunwaptan) agnostoid arthropods Kormagnostella Romanenko and Biceragnostus Ergaliev in North America. Journal of Paleontology, 87, 804–817.



2012 Pseudoolenoides

Adrain, J.M., McAdams, N.E.B. & Karim, T.S. (2012) The Middle Ordovician (Dapingian and Darriwilian) bathyurid trilobite Pseudoolenoides. Zootaxa, 3467, 1–97.



2012 Aponileus small pub

Adrain, J.M. & McAdams, N.E.B. (2012) The Lower Ordovician (upper Floian) bathyurid trilobite Aponileus Hu, with species from Utah, Texas, and Greenland. Zootaxa, 3293, 1–67.


2012 Bumatoides small pub

Carlucci, J., Westrop, S.R., Amati, L., Adrain, J.M. & Swisher, R.E. (2012) A systematic revision of the Upper Ordovician trilobite genus Bumastoides (Trilobita: Illaenidae) with new species from Oklahoma, Virginia, and Missouri. Journal of Systematic Palaeontology, 10, 679–723.



2011 Randaynia

Landing, E., Adrain, J.M., Westrop, S.R. & Kröger, B. (2012) Tribes Hill-Rochdale Formations in east Laurentia: proxies for Early Ordovician (Tremadoc) eustasy on a tropical passive margin (New York and west Vermont). Geological Magazine, 149, 93–123.



2011 Trilo Class small pub

Adrain, J.M. (2011) Class Trilobita Walch, 1771. In: Zhang, Z.-Q. (ed.), Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148, 104–109.



2011 Protopliomerella small pub

McAdams, N.E.B. & Adrain, J.M. (2011) Revision of the Lower Ordovician (lower Floian; Tulean) pliomerid trilobite Protopliomerella, with new species from the Great Basin, western USA. Zootaxa, 3144, 1-113.



2011 Gladiatoria small pub

Adrain, J.M., McAdams, N.E.B. & Westrop, S.R. (2011) Affinities of the Lower Ordovician (Tulean; lower Floian) trilobite Gladiatoria, with species from the Great Basin, western United States. Memoirs of the Association of Australasian Palaeontologists, 42, 321-367.



2011 Psalikilopsis small pub

Adrain, J.M., McAdams, N.E.B., Westrop, S.R. & Karim, T. S. (2011) Systematics and affinity of the Lower Ordovician (Tulean; lower Floian) trilobite Psalikilopsis. Memoirs of the Association of Australasian Palaeontologists, 42, 369-416.



2011 ISOS small pub

Adrain, J.M. (2011) A new trilobite biostratigraphy for the Lower Ordovician of western Laurentia and prospects for international correlation using pelagic trilobites. In: Gutiérrez-Marco, J. C., Rábano, I., Garcia-Bellido, D. (Eds.), Ordovician of the World. Cuadernos del Museo Geominero, 14, 41-42.



Landing et al 2011 small pub

Landing, E., Westrop, S.R. & Adrain, J. M. (2011) The Lawsonian Stage - the Eoconodontus notchpeakensis (Miller, 1969) FAD and HERB carbon isotope excursion define a globally correlatable terminal Cambrian stage. Bulletin of Geosciences, 86, 621-640.



2011 Lotagnostus small pub

Westrop, S.R., Adrain, J.M. & Landing, E. (2011) The Cambrian (Sunwaptan, Furongian) agnostoid arthropod Lotagnostus Whitehouse, 1936, in Laurentian and Avalonian North America: systematics and biostratigraphic significance. Bulletin of Geosciences, 86, 569-594.



2011 Panisaspis

McAdams, N.E.B. & Adrain, J.M. (2011) Panisaspis, a new genus of pliomerid trilobites from the Lower Ordovician (Ibexian; Tulean and Blackhillsian) of the Great Basin, western USA. Zootaxa, 2969, 1-68.



2011 Geol Cur small pub

Adrain, T.S., Budd, A.F., & Adrain, J.M. (2011) Digitizing collections: Experiences from the University of Iowa Paleontology Repository digitization project. Geological Curator, 9, 291-299.



2011 Hintzeia

McAdams, N.E.B., & Adrain, J.M. (2011) Systematics of the Lower Ordovician pliomerid trilobite Hintzeia, with species from the Great Basin, western USA. Zootaxa, 2910, 1-45.



2010 Westrop et al dokes smalls pub copy

Westrop, S.R., Waskiewicz, R.A., & Adrain, J.M. (2010) Systematics of Dokimocephalus and related trilobites from the Late Cambrian (Steptoean; Millardan and Furongian series) of Laurentian North America. Journal of Systematic Palaeontology, 8, 545-606.



McAdams, N.E.B., & Adrain, J. M. (2010) A new species of the Lower Ordovician pliomerid trilobite Pseudocybele and its biostratigraphic significance. Zootaxa, 2550, 21-38.



Adrain, J.M., McAdams, N.E.B. & Westrop, S.R. (2009) Trilobite biostratigraphy and revised bases of the Tulean and Blackhillsian Stages of the Ibexian Series, Lower Ordovician, western United States. Memoirs of the Association of Australasian Palaeontologists, 37, 5435-58.



2010 Heckethornia

McAdams, N.E.B. & Adrain, J.M. (2009) Heckethornia, a new genus of dimeropygid trilobites from the Lower Ordovician (Ibexian; Tulean and Blackhillsian) of the Great Basin, western USA. Canadian Journal of Earth Sciences, 46, 875-914.



McAdams, N.E.B. & Adrain, J.M. (2009) New pliomerid trilobite genus Lemureops from the Lower Ordovician (Ibexian; Tulean, Blackhillsian) of western Utah, USA. Memoirs of the Association of Australasian Palaeontologists, 37, 491-540.



2009 Xenocheilos small pub

Westrop, S.R. & Adrain, J.M. (2009) The late Cambrian (Furongian; Steptoean) trilobite genus Xenocheilos Wilson, 1949: systematics and biostratigraphic significance. Memoirs of the Association of Australasian Palaeontologists, 37, 351-368.



Adrain, J.M., Peters, S.E. & Westrop, S.R. (2009) The Marjuman trilobite Cedarina Lochman: thoracic morphology, systematics, and new species from western Utah and eastern Nevada, USA. Zootaxa, 2218, 35-58.



Adrain, J.M. (2009) New and revised species of the aulacopleurid trilobite Maurotarion from the Lower Devonian (Pragian) of Nevada. Zootaxa, 2215, 1-23.



Westrop, S.R. & Adrain, J.M. (2009) The Late Cambrian (Steptoean; Furongian) trilobite Pseudokingstonia Palmer, 1965 in North America. Canadian Journal of Earth Sciences, 46, 355-360.



Adrain, J.M., Chatterton, B.D.E. & Kloc, G.J. (2008) Systematics of the koneprusiine trilobites, with new taxa from the Silurian and Devonian of Laurentia. Journal of Paleontology, 82, 657-675.



Westrop, S.R., Eoff, J.D., Ng, T.-W., Dengler, A.A. & Adrain, J.M. (2008) Classification of the Late Cambrian (Steptoean) trilobite genera Cheilocephalus Berkey, 1898 and Oligometopus Resser, 1936 from Laurentia. Canadian Journal of Earth Sciences, 45, 725-744.



Adrain, J.M. (2008) A global species database of Trilobita: progress, results, and revision of the Treatise. In: Rábano, I., Gozalo, R. & Garcia-Bellido, D. (Eds.), Advances in trilobite research. Cuadernos del Museo Geominero, 9, 27-28.




McAdams, N.E.B. & Adrain, J.M. (2008) A new genus of dimeropygid trilobites from the Lower Ordovician (Ibexian) of western Utah, USA. In: Rábano, I. & Garcia-Bellido, D. (Eds.), A new genus of dimeropygid trilobites from the Lower Ordovician (Ibexian) of western Utah, USA. Cuadernos del Museo Geominero, 9, 251-252.



Adrain, J.M. & Westrop, S.R. (2007) Bearriverops, a new Lower Ordovician trilobite genus from the Great Basin, western USA, and classification of the family Dimeropygidae. Canadian Journal of Earth Sciences, 44, 337-366.



Adrain, J.M. & Westrop, S.R. (2007) The hystricurid trilobite Metabowmania in the Lower Ordovician (Ibexian; Stairsian) of the Great Basin, Utah and Idaho, USA. Memoirs of the Association of Australasian Palaeontologists, 34, 227-242.



Westrop, S.R. & Adrain, J.M. (2007) Bartonaspis new genus, a trilobite species complex from the base of the Upper Cambrian Sunwaptan Stage in North America. Canadian Journal of Earth Sciences, 44, 987-1003.



Westrop, S.R., Waskiewicz, R.A. & Adrain, J.M. (2007) The Late Cambrian (Steptoean) trilobite genus Bynumina Resser, 1942, in North America. Memoirs of the Association of Australasian Palaeontologists, 34, 357-376.



Adrain, J.M. & Tetreault, D.K. (2006) The brachymetopid trilobite Radnoria in the Silurian (Wenlock) of New York State and Arctic Canada. Canadian Journal of Earth Sciences, 42, 2087-2096. (for 2005)



Adrain, J.M. & Westrop, S.R. (2006) New genus of dimeropygid trilobites from the earliest Ordovician of Laurentia. Acta Palaeontologica Polonica, 51, 541-550.



Adrain, J.M. & Westrop, S.R. (2006) Notchpeakia, a new genus of Upper Cambrian (Sunwaptan) "entomaspidid" trilobites. Journal of Paleontology, 80, 1152-1171.



Adrain, J.M. & Westrop, S.R. (2006) New earliest Ordovician trilobite genus Millardicurus: The oldest known hystricurid. Journal of Paleontology, 80, 650-671.



Adrain, J.M. (2005) Aulacopleurid trilobites from the Upper Ordovician of Virginia. Journal of Paleontology, 79, 542-563.



Adrain, J.M. & Westrop, S.R. (2005) Lower Ordovician trilobites from the Baumann Fiord Formation, Ellesmere Island, Arctic Canada. Canadian Journal of Earth Sciences, 42, 1523-1546.



Adrain, J.M. & Westrop, S.R. (2005) Late Cambrian ptychaspidid trilobites from western Utah: implications for trilobite systematics and biostratigraphy. Geological Magazine, 142, 377-398.



Adrain, J.M. & Westrop, S.R. (2004) A Late Cambrian (Sunwaptan) silicified trilobite fauna from Nevada. Bulletins of American Paleontology, 365, 1-51.




Adrain, J.M., Edgecombe, G.D., Fortey, R.A., Laurie, J., McCormick, T., Owen, A.W., Waisfeld, B.G., Webby, B.D., Westrop, S.R. & Zhou, Z.-Y. (2004) Trilobites. In: Webby, B.D., Droser, M.L. & Paris, F. (Eds.), The Great Ordovician Biodiversification Event. Columbia University Press, New York, pp. 231-254.



Adrain, J.M., Lee, D.-C., Westrop, S.R., Chatterton, B.D.E. & Landing, E. (2003) Classification of the trilobite subfamilies Hystricurinae and Hintzecurinae subfam. nov., with new genera from the Lower Ordovician (Ibexian) of Idaho and Utah. Memoirs of the Queensland Museum, 48, 553-586.



Jell, P.A. & Adrain, J.M. (2003) Available generic names for trilobites. Memoirs of the Queensland Museum, 48, 331-553.



Adrain, J.M. & Westrop, S.R. (2003) Paleobiodiversity: We need new data. Paleobiology, 29, 22-25.



Adrain, J.M. (2003) Validity and composition of the Silurian trilobite genera Borealarges and Dicranogmus, with new species from the Canadian Arctic. Canadian Journal of Earth Sciences, 40, 749-763.


Adrain, J.M., Westrop, S.R., Landing, E. & Fortey, R.A. (2001) Systematics of the Ordovician trilobites Ischyrotoma and Dimeropygiella, with species from the type Ibexian area, western U.S.A. Journal of Paleontology, 75, 947-971.



Adrain, J.M. & Westrop, S.R. (2001) Stratigraphy, phylogeny, and species sampling in time and space. In: Adrain, J.M., Edgecombe, G.D. & Lieberman, B.S. (Eds.), Fossils, Phylogeny, and Form: An Analytical Approach. Kluwer, New York, pp. 291-322.



Adrain, J.M. (2001) Systematic paleontology. Journal of Paleontology, 75, 1055-1057.




Westrop, S.R. & Adrain, J.M. (2001) Sampling at the species level: impact of spatial biases on diversity gradients. Geology, 29, 903-906.


Brochu, C.A., Bryant, H.N., Theodor, J.M., O'Leary, M.A., Adrain, J.M. & Sumrall, C.D. (2001) Modern phylogenetics in paleontology: comments on Vermeij, 1999. Paleobiology, 27, 174-176.



Adrain, J.M. & Westrop, S.R. (2000) An empirical assessment of taxic paleobiology. Science, 289, 110-112.



Adrain, J.M., Westrop, S.R., Chatterton, B.D.E. & Ramsköld, L. (2000) Silurian trilobite alpha diversity and the end-Ordovician mass extinction. Paleobiology, 26, 625-646.



Adrain, J.M. & Westrop, S.R. (1999) Trilobite paleobiology: Past, present, and future. Journal of Paleontology, 73, 161-163.



Hughes, N.C., Chapman, R.E. & Adrain, J.M. (1999) The stability of thoracic segmentation in trilobites: a case study in developmental and ecological constraints. Evolution and Development, 1, 24-35.



Adrain, J.M., Fortey, R.A. & Westrop, S.R. (1998) Post-Cambrian trilobite diversity and evolutionary faunas. Science, 280, 1922-1925.




Adrain, J.M. (1998) Systematics of the Acanthoparyphinae (Trilobita), with species from the Silurian of Arctic Canada. Journal of Paleontology, 72, 698-718.



Westrop, S.R. & Adrain, J.M. (1998) Trilobite diversity and the Ordovician Radiation: a reply to Miller et al. Paleobiology, 24, 529-533.


Westrop, S.R. & Adrain, J.M. (1998) Trilobite alpha diversity and the reorganization of Ordovician benthic marine communities. Paleobiology, 24, 1-16.



Davidek, K., Landing, E., Bowring, S.A., Westrop, S.R., Rushton, A.W.A., Fortey, R.A. & Adrain, J.M. (1998) New uppermost Cambrian U-Pb date from Avalonian Wales and age of the Cambrian-Ordovician boundary. Geological Magazine, 135, 305-309.



Adrain, J.M. (1997) Proetid trilobites from the Silurian (Wenlock-Ludlow) of the Cape Phillips Formation, Canadian Arctic Archipelago. Palaeontographia Italica, 84, 21-111.



Adrain, J.M. & Fortey, R.A. (1997) Ordovician trilobites from the Tourmakeady Limestone, western Ireland. Bulletin of the Natural History Museum, London, Geology Series, 53, 79-115.



Adrain, J.M. & Ramsköld, L. (1997) Silurian Odontopleurinae (Trilobita) from the Cape Phillips Formation, Arctic Canada. Journal of Paleontology, 71, 237-261.



Adrain, J.M. & Edgecombe, G.D. (1997) Silurian (Wenlock) calymenid trilobites from the Cape Phillips Formation, central Canadian Arctic. Journal of Paleontology, 71, 657-682.


Adrain, J.M. & Edgecombe, G.D. (1997) Silurian encrinurine trilobites from the central Canadian Arctic. Palaeontographica Canadiana, 14, 1-109.



Adrain, J.M. & Kloc, G.J. (1997) Lower Devonian aulacopleuroidean trilobites from Oklahoma. Journal of Paleontology, 71, 703-712.



Adrain, J.M. (1996) A new otarionine trilobite from the Henryhouse Formation (Silurian, Ludlow) of Oklahoma. Journal of Paleontology, 70, 611-614.



Adrain, J.M. & Ramsköld, L. (1996) The lichid trilobite Radiolichas in the Silurian of Arctic Canada and Gotland, Sweden. Geological Magazine, 133, 147-158.



Adrain, J.M. & MacDonald, E.W. (1996) Phacopid trilobites from the Silurian of Arctic Canada. Journal of Paleontology, 70, 1091-1094.



Adrain, J.M. & Chatterton, B.D.E. (1996) The otarionine trilobite Cyphaspis, with new species from the Silurian of northwestern Canada. Journal of Paleontology, 70, 100-110.



Adrain, J.M. & Edgecombe, G.D. (1996) Devonian aulacopleurid trilobites of the Malvinokaffric Realm. Geobios, 29, 417-436.



Adrain, J.M., Chatterton, B.D.E. & Blodgett, R.B. (1995) Silurian trilobites from southwestern Alaska. Journal of Paleontology, 69, 723-736.


Adrain, J.M. & Edgecombe, G.D. (1995) Balizoma and the new genera Aegrotocatellus and Perirehaedulus: encrinurid trilobites from the Douro Formation (Silurian, Ludlow) of the central Canadian Arctic. Journal of Paleontology, 69, 736-752.



Adrain, J.M. & Chatterton, B.D.E. (1995) The otarionine trilobites Harpidella and Maurotarion, with species from northwestern Canada, the United States, and Australia. Journal of Paleontology, 69, 307-326.



Adrain, J.M. & Chatterton, B.D.E. (1995) Aulacopleurine trilobites from the Llandovery of northwestern Canada. Journal of Paleontology, 69, 326-340.



Edgecombe, G.D. & Adrain, J.M. (1995) Silurian calymenid trilobites from the United States. Palaeontographica Abteilung A, 235, 1-19.



Adrain, J.M. (1994) The lichid trilobite Borealarges n. gen., with species from the Silurian of Arctic Canada. Journal of Paleontology, 68, 1081-1099.



Adrain, J.M. & Chatterton, B.D.E. (1994) The aulacopleurid trilobite Otarion, with new species from the Silurian of northwestern Canada. Journal of Paleontology, 68, 305-323.



Adrain, J.M. & Wilson, M.V.H. (1994) Early Devonian cephalaspids (Vertebrata: Osteostraci: Cornuata) from the southern Mackenzie Mountains, N.W.T., Canada. Journal of Vertebrate Paleontology, 14, 301-319.


Ramsköld, L., Adrain, J.M., Edgecombe, G.D. & Siveter, D.J. (1994) Silurian calymenid trilobite Alcymene n. gen., with new species from the Ludlow of Gotland, Sweden. Journal of Paleontology, 68, 556-569.


Adrain, J.M. & Chatterton, B.D.E. (1993) A new rorringtoniid trilobite from the Ludlow of Arctic Canada. Canadian Journal of Earth Sciences, 30, 1634-1643.


Adrain, J.M. (1992) Machaeridian classification. Alcheringa, 16, 15-32.



Adrain, J.M., Chatterton, B.D.E. & Cocks, L.R.M. (1991) A new species of machaeridian from the Silurian of Podolia, USSR, with a review of the Turrilepadidae. Palaeontology, 34, 637-651.


Adrain, J.M. & Chatterton, B.D.E. (1990) Odontopleura (Trilobita, Silurian), and a method of constrained congruency analysis. Journal of Paleontology, 64, 600-614.